Leon studied the anatomy and functional morphology of theropod dinosaur gastralia with Phil at the Royal Tyrrell Museum of Palaeontology in Drumheller from 1995–1996. This was associated with Leon’s master’s research at Utrecht University, Netherlands around the same time. His work on dinosaur gastralia and potential respiratory functions has led to broader research on aspiration systems in modern-day crocodylians and birds. Leon received a PhD in Biology at Harvard University, and is now Professor of Vertebrate Paleontology and Evolution at Maastricht University, Netherlands.
Claessens, L.P.A.M. 2004. Dinosaur gastralia; origin, morphology, and function. Journal of Vertebrate Paleontology 24(1): 89–106. DOI: 10.1671/A1116-8