Peer-Reviewed Journal Publications (Google Scholar Link)

Normandeau, J., S.J. Kutz, M. Hebblewhite,E.H. Merrill (2020). Living with liver flukes: Does migration matter? Internatl. J. Parasitology: Parasites and Wildlife. 12:76-84.

Merkle, J.A., N.J. Anderson, D.L. Baxley, M. Chopp, L.C. Gigliotti, J.A. Gude, T.M. Harms, H.E. Johnson,E.H. Merrill, M.S. Mitchell, T.W. Mong, J. Nelson, A.S. Norton, M.J. Sheriff, E. Tomasik, K.R. Vanbeek, (2019) A collaborative approach to bridging the gap between wildlife managers and researchers. J. Wildl. Manage. 83:1644-1651. doi: 10.1002/jwmg.21759

Smolko, P, D.Seidel, E. Merrill, M. Pybus, A. Hubbs, M. Ball (2019) Inferring the effects of deer population and landscape structure on spread of the chronic wasting disease in Alberta. PRION. 13:9-1.

Muhly, T.B.; C.A. Johnson, M. Hebblewhite, E.W. Neilson, D. Fortin, J.M. Fryxell, A.D.M. Latham, M.C. Latham, P.D. McLoughlin, E.H. Merrill, P.C. Paquet, B.R. Patterson, F. Schmiegelow, M. Musiani, (2019). Functional response of wolves to human development across boreal North America. Ecol. Evol. 18:‏ 10801-10815. 

Berg, Jodi E., M. Hebblewhite, C.C. St Clair,E.H. Merrill (2019) prevalence and mechanisms of partial migration in ungulates. Front. Ecol. Evol. 7:324 doi: 10.3389/fevo.2019.00325

Normandeau, J., K. MacAulay, J. Berg and E. Merrill (2019).  Identifying guard hairs of Rocky Mountain carnivores. Wildlife Society Bulletin: 42:706-712.

Visscher, D.R.,E.H. Merrill (2018) Functional connectivity in ruminants: A generalized state-dependent modelling approach. PLoS One. 13(6): 22

Hebblewhite M., D.R. Eacker, S. Eggeman, H. Bohm, E.H. Merrill (2018) Density-independent predation affects migrants and residents equally in a declining partially migratory elk population. doi:10.1111/oik.05304 (epub)

Smolko, Peter, R.Kropil, T. Pataky, A. Veselovska, E.H. Merrill (2018) Why do migrants move downhill? The effects of increasing predation and density on red deer altitudinal migration in temperate Carpathian forests. Mammal Research. 63(3): 297-305

Tucker, Marlee A.; Böhning-Gaese, Katrin; Fagan, William F.; Fryxell, John M.; Moorter, Bram Van; Alberts, Susan C.; Ali, Abdullahi H.; Allen, Andrew M.; Attias, Nina; Avgar, Tal; Bartlam-Brooks, Hattie; Bayarbaatar, Buuveibaatar; Belant, Jerrold L.; Beyer, Dean; Bidner, Laura; Van Beest, Floris M.; Blake, Stephen; Blaum, Niels; Bracis, Chloe; Brown, Danielle; De Bruyn, PJ Nico; Cagnacci, Francesca; Calabrese, Justn M.; Camilo-Alves, Constanca; Chamaille-Jammes, Simon; Chiaradia, Andre; Davidson, Sarah C.; Dennis, Todd; DeStefano, Stephen; Diefenbach, Duane; Douglas-Hamilton, Iain; Fennessy, Julian; Fichtel, Claudia; Fiedler, Wolfgang; Fischer, Christina; Fischhoff, Ilya; Fleming, Christian H.; Ford, Adam T.; Fritz, Susanne A.; Gehr, Benedikt; Goheen, Jacob R.; Gurarie, Eliezer; Hebblewhite, Mark; Heurich, Marco; Hewison, AJ Mark; Hof, Christian; Hurme, Edward; Isbell, Lynne A.; Janssen, Rene; Jeltsch, Florian; Kaczensky, Petra; Kane, Adam; Kappeler, Peter M.; Kauffman, Matthew; Kays, Roland; Kimuyu, Duncan; Koch, Flavia; Kranstauber, Bart; LaPoint, Scott; Leimgruber, Peter; Linnell, John DC; Lopez-Lopez, Pascual; Markham, A. Catherine; Mattisson, Jenny; Medici, Emilia Patricia; Mellone, Ugo; Merrill, Evelyn; De Miranda Mourao, Guilherme; Morato, Ronald G.; Morellet, Ronaldo G.; Morellet, Nicolas; Morrison, Thomas A.; Diaz-Munoz, Samuel L.; Mysterud, Atle; Nandintsetseg, Dejid; Nathan, Ran; Niamir, Aidin; Odden, John; O’Hara, Robert B.; Oliveira-Santos, Luis Gustavo R.; Olson, Kirk A.; Patterson, Bruce D.; Cunha de Paula, Rogerio; Pedrotti, Luca; Reineking, Bjorn; Rimmler, Martin; Rogers, Tracey L.; Rolandsen, Christer Moe; Rosenberry, Christopher S.; Rubenstein, Daniel I.; Safi, Kamran; Said, Sonia; Sapir, Nir; Sawyer, Hall; Schmidt, Niels Martin; Selva, Nuria; Sergiel, Agnieszka; Shiilegdamba, Enkhtuvshin; Silva, Joao Paulo; Singh, Navinder; Solberg, Erling J.; Spiegel, Orr; Strand, Olav; Sundaresan, Siva; Ullmann, Wiebke; Voigt, Ulrich; Wall, Jake; Wattles, David; Wikelski, Martin; Wilmers, Christopher C.; Wilson, John W.; Wittemyer, George; Filip, Zieba; Zwijacz-Kozica, Tomasz; Mueller, Thomas (2018) Moving in the Anthropocene: Global reductions in terrestrial mammalian movements 359(6374): 466-469

Geary, A.B, E.H. Merrill, J.G. Cook, R.C. Cook, and L.L. Irwin. Resubmitted. Elk nutritional resources: Herbicides, herbivory and forest succession at Mount St. Helens. Forest Ecology and Management.

Schlägel, U.E., E.H. Merrill, and M. A. Lewis. In press. Territory surveillance and prey management: wolves keep track of space and time. Ecology and Evolution.

Visscher, D. and E. H. Merrill. 2017. Hierarchical trade-offs between risk and reward mediated by behavior. Mammal Research 62:129-140. 

Nobert, B. E. H. Merrill, M. Pybus, T. Bollinger, and Y. T. Hwang. 2016. Landscape connectivity predicts chronic wasting disease risk in Canada. Journal of Applied Ecology 53: 1450-1459.

Cagnacci, F., S. Focardi, A. Ghisla, B. van Moorter, E. H. Merrill, E. Gurarie, M. Heurich, A. Mysterud, J. Linnell, M. Panzacchi, R. May, T. Nygard, C. Rolandsen, and M. Hebblewhite. 2016. How many routes lead to migration? Comparison of methods to assess and characterize migratory movements. Journal of Applied Ecology 85: 54-68.

Eggeman, S. L., M. Hebblewhite, H. Bohm, J. Whittington, and E. H. Merrill. 2016. Behavioural flexibility in migratory behaviour in a long-lived large herbivore. Journal of Animal Ecology 85: 785-797.

Potapov, A., E. H. Merrill, M. Pybus, M. A. Lewis. 2016. Chronic Wasting Disease: Transmission mechanisms and the possibility of harvest management. PLOS ONE 11:e0151039. 

Boyce, M. S., C. J. Johnson, E. H. Merrill, S. E. Nielsen, E. J. Solberg and B. van Moorter. 2015. Can habitat selection predict abundance? Journal of Animal Ecology 85: 11-20.

Lele, S. R., E. H. Merrill, J.Keim and M.S. Boyce. 2013. Selection, use, choice and occupancy: clarifying concepts in resource selection studies. Journal of Animal Ecology 82: 1183-1191.

Robinson, B. G and E. H.Merrill. 2013. Foraging-vigilance trade-offs in a partially migratory population: comparing migrants and residents on a sympatric range. Animal Behaviour 85: 849-856.

Potapov, A., E. H. Merrill, M. Pybus, D. Coltman, and M. Lewis. 2012. Chronic wasting disease: possible transmission mechanisms in deer. Ecological Modelling 250: 244-257.

Habib, T. J., D. A. Moore, and E. H. Merrill. 2012. Detection and stratification approaches for aerial surveys of deer in prairie parklands. Wildlife Research 39: 593-602.

Webb, N. F. and E. H. Merrill. 2012. Simulating carnivore movements: an occupancy-abundance relationship for surveying wolves. Wildlife Society Bulletin 36: 240-247.

Potapov, A., E. H. Merrill, M. A. Lewis. 2012. Wildlife disease elimination and density dependence. Proceedings Royal Society B. 279: 3139-3145.

McPhee, H. M, N. F. Webb, and E. H. Merrill. 2012. Hierarchical predation: wolf (Canis lupus) selection along hunt paths and at kill sites. Canadian Journal Zoology 90:555-563.

Mckenzie, H.W., E. H. Merrill, R. J. Spiteri, and M.A. Lewis. 2012. How linear features alter predator movement and the functional response. Interface Focus 2:205-216. 

Rees, E. E., E. H. Merrill, T. K. Bollinger, Y. T. Hwang, M. Pybus, D. Coltman. 2012. Targeting the detection of chronic wasting disease using the hunter harvest during early phases of an outbreak in Saskatchewan, Canada. Preventive Veterinary Medicine 104:149-159.

McPhee, H.M., N.F. Webb, E.H. Merrill. 2012. Time-to-kill: measuring attack rates in a heterogeneous landscape with multiple prey types. Oikos 121:711-720. 

Hebblewhite, M. and E.H. Merrill. 2012. Demographic balancing of migrant and resident elk in a partially migratory population through forage–predation tradeoffs. Oikos 120-1860-1870. 

Nelson, B., M. Hebblewhite, V. Ezenwa, T. Shury, E.H. Merrill, P.C. Paquet, F. Schmiegelow, D. Seip, G. Skinner, N. Webb. 2012. Prevalence of antibodies to canine parvovirus and distemper virus in wolves in the canadian rocky mountains. Journal Wildlife Diseases 48:68-76.

Webb, N. F., J.A. Allen and E. H. Merrill. 2011. Demography of a harvested wolf population in west-central Alberta, Canada. Canadian Journal Zoology 89:744-752. 

Robinson, B. G. and E. H. Merrill. 2011. The influence of snow on the functional response of grazing ungulates. Oikos 164:265-275.

Romme, W.H., M. . Boyce, R. Gresswell, E. H. Merrill, G. W. Minshall,C.Whitlock, and M. G.Turner. 2011. Twenty years after the 1988 Yellowstone fires: Lessons about disturbance and ecosystems. Ecosystems 14:1196. doi:10.1007/s10021-011-9470-6. 

Habib, T. J., E. H. Merrill, M. J. Pybus, and D. W. Coltman. 2011. Modelling landscape effects on density-contact rate relationships of deer in eastern Alberta: implications for chronic wasting disease. Ecological Modeling 222:2722-2732. 

Cullingham C.I., E.H. Merrill, M. Pybus, Bollinger, TK., G.D., Wilson, D.W. Coltman. 2011. Broad and fine-scale genetic analysis of white-tailed deer populations: estimating the relative risk of chronic wasting disease spread. Evolutionary Applications 4:116-131. 

Cullingham, C.I., S.M. Nakada, E.H. Merrill, T.K. Bollinger, M.J. Pybus, and D.W. Coltman. 2011. Multiscale population genetic analysis of mule deer (Odocoileus hemionus hemionus) in western Canada sheds new light on the spread of chronic wasting disease. Canadian Journal Zoology 89:134-147. 

Merrill, E., H. Sand, B. Zimmermann, H. McPhee, N. Webb, M. Hebblewhite, P. Wabakken and J. Frair. 2010. Building a mechanistic understanding of predation with GPS-based movement data. Philosophical Transactions Royal Society B 365:2279-2288. 

Robinson, B.G., M. Hebblewhite, E.H. Merrill. 2010. Are migrant and resident elk (Cervus elaphus) exposed to similar forage and predation risk on their sympatric winter range? Oecologia 164:265-275.

Morales, J. P.R. Moorcroft, J. Matthiopoulos, J.L. Frair, J.G. Kie, R.A. Powell, E. H. Merrill, and D.T. Haydon. 2010. Building the bridge between animal movements and population dynamics. Philosophical Transactions Royal Society B 365:2289-2301. 

Owen-Smith, N., J. M. Fryxell, and E. H. Merrill. 2010. Foraging theory upscaled: the behavioural ecology of herbivore movement. Philosophical Transactions Royal Society B 365:2267-2278.

Van Moorter B., D.R. Visscher, C.L. Jerde, J.L. Frair and E.H. Merrill. 2010. Identifying movement states from location data using cluster analysis. Journal Wildlife Management 74:588-594. 

McInenly, L.E., E.H. Merrill, J.F. Cahill, and N.G. Juma 2010. Festuca campestris alters root morphology and growth in response to simulated grazing and nitrogen form. Functional Ecology 24:283-292. 

Hebblewhite, M. and E.H. Merrill. 2009. Trade-offs between predation risk and forage differ between migrant strategies in a migratory ungulate. Ecology 90:3445-3454.

Wilson, G.A., S.M. Nakada, T. K. Bollinger, M.J. Pybus, E.H. Merrill, and D.W. Coltman. 2009. Polymorphisms at the PRNP Polymorphisms at the PRNP gene influence susceptibility to chronic wasting disease in two species of deer to chronic wasting disease in two species of deer (Odocoileus spp.) in western Canada. Journal Toxicology and Environmental Health 72:1025-1029.

Hebblewhite, M., R.H. Munroe, and E.H. Merrill. 2009. Trophic consequences of postfire logging in a wolf-ungulate system. Forest Ecology and Management 257:1053-1062.

McKenzie H, M. Lewis, and E. Merrill. 2009. First passage time of animal movement and insights in to the functional response. Bulletin of Mathematical Biology 71:107-129.

Visscher, D. and E.H. Merrill. 2009. Temporal dynamics of forage succession for ungulates at two scales: Implications for forest management. Forest Ecology and Management 257:96-106.

Wolf, M. J.L. Frair, E.H. Merrill, and P. Turchin. 2009. The attraction of the unknown:the importance of spatial familiarity in habitat selection in wapiti (Cervus elaphus). Ecography 32:401-410.

McKenzie, H.W., C.L. Jerde, D.R. Visscher, E.H. Merrill, and M.A. Lewis. 2008. Inferring linear feature use in the presence of GPS measurement error. Environmental and Ecological Statistics 16:531-546.

Frair, J.L., E.H. Merrill, H.L. Beyer, and J.M. Morales. 2008. Thresholds in landscape connectivity and mortality risks in response to growing road networks. Journal of Applied Ecology 45:1504-1513. (Editor`s recommended article)

Allen, J.R., L.E. McInenly, E.H. Merrill, and M.S. Boyce. 2008. Using resource selection functions to improve estimation of elk population numbers. Journal of Wildlife Management 72:1798-1804.

Webb, N.F., M. Hebblewhite, and E.H. Merrill. 2008. Statistical methods for identifying wolf kill sites using global positioning system locations. Journal Wildlife Management 72:798-807.

Hebblewhite, M., E.H. Merrill, and G. McDermid. 2008. A multi-scale test of the forage maturation hypothesis in a partially migratory ungulate population. Ecological Monographs 78:141-166. (Cover photo)

Hebblewhite, M. and E.H. Merrill. 2008. Modeling wildlife-human relationships for social species with mixed-effects resource selection models. Journal Applied Ecology 45:834-844. (Cover photo)

Beyer, H.L., E.H. Merrill, N. Varley, and M.S. Boyce. 2007. Willow on Yellowstone National Park’s northern range: evidence for a trophic cascade in a large mammalian predator-prey system. Ecological Applications 17:1563-1571.

Hebblewhite, M., M. Percy, and E.H. Merrill. 2007. Are all GPS collars created equal? Correcting habitat-induced bias using three brands in the Central Canadian Rockies. Wildlife Society Bulletin 71:2026-2033.

Hebblewhite, M. and E.H. Merrill. 2007. Multi-scale wolf predation risk for elk: does migration reduce risk. Oecologia 152:377-387.

Koper, N., F.K.A. Schmiegelow, and E.H. Merrill. 2007. Residuals cannot distinguish between ecological effects of habitat amount and fragmentation: implications for the debate. Landscape Ecology 22:811-820. (Honorable mention from Landscape Ecology: 2008)

Frair, J.L., E.H. Merrill, J.W. Allen, and M.S. Boyce. 2007. Know thy enemy: lessons from elk translocations into risky landscapes. Journal of Wildlife Management 71:541-554.

Hebblewhite, M., E.H. Merrill, L.E. Morgantini, C.A. White, J.R. Allen, E. Bruns, L. Thurston, and T.E. Hurd. 2006. Is the migratory behaviour of montane elk (Cervus elaphus) herds in peril? The case of Alberta’s Ya Ha Tinda elk herd. Wildlife Society Bulletin 34:1280-1294.

Johnson, C.J., S.E. Nielsen, E.H. Merrill, T.L. McDonald, and M.S. Boyce. 2006. Resource selection functions based on use-availability data: theoretical motivation and evaluation methods. Journal Wildlife Management 70:347-357.

Visscher, D.R., E.H. Merrill, D. Fortin, and J.L. Frair. 2006. Estimating woody browse availability for ungulates at increasing snow depths. Forest Ecology and Management 222:348-354.

Mao, J.S., M.S. Boyce, D.W. Smith, F.J. Singer, D.J. Vales, and E.H. Merrill. 2005. Habitat selection by elk before and after wolf reintroduction in Yellowstone National Park. Journal of Wildlife Management 69:1691-1707.

Anderson, D.P., J.D. Forester, M.G. Turner, J.L. Frair, E.H. Merrill, D. Fortin, J. Mao, and M.S. Boyce. 2005. Factors influencing home range sizes of elk (Cervus elaphus) in North American landscapes. Landscape Ecology 20:257-271.

Frair, J.L., E.H. Merrill, D.R. Visscher, D. Fortin, H.L. Beyer, and J.M. Morales. 2005. Scales of movement by elk (Cervus elaphus) in response to heterogeneity and predation risk. Landscape Ecology 20:273-287.

Hebblewhite, M., E.H. Merrill, and T.L. McDonald. 2005. Spatial decomposition of predation risk using resource selection functions: an example in a wolf-elk predator-prey system. Oikos 111:101-111.

Merrill, E.H., J.J. Hartigan, and M.W. Meyer. 2005. Does prey biomass or mercury exposure affect loon chick survival in Wisconsin? Journal Wildlife Management 69:57-67.

Morales, J.M., D. Fortin, J.L. Frair, and E.H. Merrill. 2005. Adaptive models for large herbivore movements in heterogeneous landscapes. Landscape Ecology 20:301-316.

Fortin. D., M.S. Boyce, E.H. Merrill, and J.M. Fryxell. 2004. Foraging costs of vigilance in large mammalian herbivores. Oikos 106:1-9.

Fortin. D., M.S. Boyce, and E.H. Merrill. 2004. Multi-tasking by mammalian herbivores: overlapping processes during foraging. Ecology 85:2312-2322.

Frair, J.L., S.E. Nielson, E.H. Merrill, S.R. Lele, M.S. Boyce, R.H. Munro, G.B. Stenhouse, and H.L. Beyer. 2004. Approaches for removing GPS-collar bias in habitat-selection studies. Journal of Applied Ecology 41:201-212.

Boyce, M.S., J. Mao, E.H. Merrill, D. Fortin, M.G. Turner, J. Fryxell, and P. Turchin. 2003. Scale and heterogeneity in habitat selection by elk in Yellowstone National Park. EcoScience 10:421-431.

Merrill, E.H., H.L. Beyer, G.P. Jones, and G.W. McDaniel. 2003. Deciduous woodland conservation under heavy deer browsing on Devils Tower National Monument, USA. Journal for Nature Conservation 10:221-232.

Merrill, E.H. and P.J.S. Coleberg. 2003. Defoliation, waterlogging and dung influences allocation patterns of Descampsia caespitosa. Journal of Range Management 56:634-639.

Driese, K.L., W.A. Reiners, E.H. Merrill, and K.G. Gerow. 1997. A digital GIS land cover map of Wyoming: a tool for vegetation analysis. Journal Vegetation Science 8:133-146.

Merrill, E.H. 1994. Summer foraging ecology of elk in the Mount St. Helens blast zone. Canadian Journal Zoology 72:303-311.

Merrill, E.H., N. Stanton, and H. Hak. 1994. Responses of bluebunch wheatgrass, Idaho fescue, and nematodes to ungulate grazing in Yellowstone National Park. Oikos 69:231-240.

Merrill, E.H., L.E. Kuck, and T.A. Hemker. 1994. Impacts of mining facilities on fall migration of mule deer. Wildlife Society Bulletin 22:68-73.

Merrill, E.H., R.W. Marrs, M.B. Brodahl, and M.S. Boyce. 1993. Estimations of green herbaceous phytomass in Yellowstone National Park using remote sensing. Journal Range Management 46:151-157.

Merrill, E.H. 1991. Thermal constraints on the use of cover types and activity patterns of elk. Applied Animal Behavior Science 29:251-267.

Boyce, M.S. and E.H. Merrill. 1991. Ungulate responses to the 1988 fires in Yellowstone National Park. Tall Timbers Fire Ecology 17:121-132.

Kuck, L.E., G.L. Hompland, and E.H. Merrill. 1985. Elk calf response to stimulated mine disturbances in Southeast Idaho. Journal Wildlife Management 49:751-757.

Merrill, E.H., H.F. Mayland, and J.M. Peek. 1982. Shrub responses after fire in an Idaho Ponderosa pine community. Journal Wildlife Management 46:496-502.

Merrill, E.H., H.F. Mayland, and J.M. Peek. 1980. Effects of wildfire on herbaceous vegetation on xeric sites in the Selway-Bitterroot Wilderness. Journal Range Management 33:363-367.

Merrill, E.H. 1978. Bear depredation at back country campgrounds in Glacier National Park. Wildlife Society Bulletin 6:123-127.

* Student names are underlined.

Published as Editor-in Chief Messages in Journal of Wildlife Management