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(Names in bold are Palmer lab members; * authorship alphabetical)
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Palmer, A.R. 2026.
DISPATCH Developmental mechanics: How elongating snake embryos coil to fit their space.
Current Biology 36:R975-R976.
(download from journal or request pdf from me)
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Palmer, A.R. 2024.
Evolution of left-right asymmetry in side-resting (pleurothetic) bivalves: A test of genes as leaders or followers.
Journal of Molluscan Studies 90:eyae040.
(download from journal or request pdf from me)
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Palmer, A.R. 2021a.
So you have to teach invertebrate biology . . .
Invertebrate Biology 140(1):e12330.
(download from journal or request pdf from me)
NEW! Play the "Guess the Invert" puzzle from the cover of the 25 Anniversary Issue of Invertebrate Biology.
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Palmer, A.R. 2021b.
Sex, camouflage, marvelous adaptations: A writing assignment that inspires.
Invertebrate Biology 140(1):e12322.
(download from journal or request pdf from me)
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Longo, S.J., W. Ray, G.M. Farley, J. Harrison, J. Jorge, T. Kaji, A.R. Palmer & S.N. Patek 2021.
Snaps of a tiny amphipod push the boundary of ultrafast, repeatable movement.
Current Biology 31:R116-R117.
(download from journal or request pdf from me)
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Miyashita, T., P. Baddam, J. Smeeton, A.P. Oel, N. Natarajan, B. Gordon, Palmer, A.R., J.G. Crump, D. Graf & W.T. Allison 2020.
nkx3.2 mutant zebrafish accommodate jaw joint loss through a phenocopy of the head shapes of Paleozoic jawless fish.
Journal of Experimental Biology 223:jeb216945.
(download from journal or request pdf from me)
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Palmer, A.R. 2019.
Obituary: Tomonari Kaji (1982-2019), evolutionary morphologist extraordinaire
Journal of Experimental Zoology (Molecular and Developmental Evolution) 332:215-218.
(download from journal or request pdf from me)
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Goto, R., I. Hirabayashi & A.R. Palmer. 2019.
Remarkably loud snaps during mouth-fighting by a sponge-dwelling worm
Current Biology 29:R603-R622.
(download from journal or request pdf from me)
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Kaji, T., C. Song, K. Murata, S. Nonaka, K. Ogawa, Y. Kondo, S. Ohtsuka & A.R. Palmer. 2019.
Evolutionary transformation of mouthparts from particle-feeding to piercing carnivory in Viper copepods: Review and 3D analyses of a key innovation using advanced imaging techniques
Frontiers in Zoology 16:35.
(download from journal, Open Access)
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(13 cites)
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Luque, J., R.M. Feldmann, H. Karasawa, C.E. Schweitzer, C.B. Cameron, K.A. Kerr, F.J. Vega, A. Duque, M. Strange, A.R. Palmer & C. Jaramillo. 2019.
Exceptional preservation of mid-Cretaceous marine arthropods and the evolution of novel forms via heterochrony
Science Advances 5:eaav3875.
(download from journal or request pdf from me)
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(7 cites)
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Miyashita, T., M.I. Coates, R. Farrar, P. Larson, P.L. Manning, R.A. Wogelius, N.P. Edwards, J. Anne, U. Bergmann, A.R. Palmer & P.J. Currie. 2019.
Hagfish from the Cretaceous Tethys Sea and a reconciliation of the morphological-molecular conflict in early vertebrate phylogeny
PNAS 116:2146-2151.
(download from journal or request pdf from me)
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Journal
Cover
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Webster, N.B. & A.R. Palmer. 2019.
How do gastropods grow synchronized shell sculpture? Effect of experimental varix manipulations on shell growth by Ceratostoma foliatum (Muricidae: Ocenebrinae)
Invertebrate Biology 138:74-88.
(download from journal or request pdf from me)
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Prado, L.A.C., Luque, J., I.S. Carvalho, A.M.F. Barreto & A.R. Palmer. 2018.
New brachyuran crabs from the Aptian-Albian Romualdo Formation, Santana Group of Brazil:
Evidence for a tethyan connection to the Araripe Basin
Acta Palaeontologica Polonica 20:160-171.
(download from journal, Open Access)
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Webster, N.B. & A.R. Palmer. 2018.
Connecting pattern to process:
How snails grow spiral shell sculpture in Nucella ostrina (Muricidae: Ocenebrinae)
Evolution & Development 20:160-171.
(download from journal or request pdf from me)
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Hakai
Story
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Newson, O., R. Basi & A.R. Palmer. 2018.
Lamellose axial shell sculpture reduces gastropod vulnerability to sea star predation
Biological Bulletin 235:24-29.
(download from journal or request pdf from me)
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Media
Resources
(14 cites)
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Kaji, T., C.S. Wirkner, A. Anker & A.R. Palmer. 2018.
Parallel saltational evolution of ultrafast movements in snapping shrimp claws
Current Biology 28:106-113.
(download from journal, Open Access)
View full sized video abstract (9min 11 sec, 150 MB) (or view smaller version on Current Biology web site)
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Luque, J., J.H. Christy, A.J.W. Hendy, M.S. Rosenberg, K.A. Kerr, R.W. Portell & A.R. Palmer. 2017.
Quaternary intertidal and supratidal crabs (Decapoda, Brachyura) from tropical America and the systematic affinities of fossil fiddler crabs
Journal of Systematic Palaeontology 16:12, 1037-1055.
(download from journal or request pdf from me)
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Journal
Cover
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Kaji, T. & A.R. Palmer. 2017.
How reversible is development? Contrast between developmentally plastic gain and loss of segments in barnacle feeding legs
Evolution 71:756-765.
(download from journal or request pdf from me)
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Palmer, A.R. 2016.
What determines direction of asymmetry: Genes, environment or chance?
Philosophical Transactions of the Royal Society of London 371:20150417.
(download from journal, Open Access)
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Kaji, T., K. Kakui, N. Miyazaki, K. Murata & A.R. Palmer. 2016.
Mesoscale morphology at nanoscale resolution: Serial block-face scanning electron microscopy reveals fine 3D detail of a novel silk spinneret system in a tube-building tanaid crustacean
Frontiers in Zoology 13:14.
(download from journal, Open Access)
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Webster, N. & A.R. Palmer. 2016.
Shaving a shell: Effect of manipulated sculpture and feeding
on shell growth and sculpture development in Nucella lamellosa (Muricidae: Ocenebrinae)
Biological Bulletin 230:1-14.
(download from journal or request pdf from me)
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Barazandeh, M. & A.R. Palmer. 2015.
Novel mating modes on wave-swept shores:
Aerial copulation and sperm release in an intertidal stalked barnacle
Marine Biology 162:881-888.
(download from journal or request pdf from me;
view video of Pollicipes mating in air)
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Palmer, A.R. 2014.
Barnacle reproduction
Access Science, McGraw Hill.
(view article or request pdf from me)
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Journal
Cover
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Miyashita, T. & A.R. Palmer. 2014.
Handed behavior in hagfish - an ancient vertebrate lineage - and a survey of lateralized behaviors in other invertebrate chordates and elongate vertebrates
Biological Bulletin 226:111-120.
(download from journal or request pdf from me)
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Barazandeh, M M., C.S. Davis & A.R. Palmer. 2014.
Where even a long penis can't help:
Evidence of long-distance spermcast mating in two acorn barnacles
Journal of Experimental Marine Biology and Ecology 454:49-54.
(download from journal or request pdf from me)
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MEDIA
COVERAGE:
Inside JEB
Science News
Echino Blog
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Hayne, K. & A.R. Palmer. 2013.
Intertidal sea stars (Pisaster ochraceus (Brandt))
alter body shape in response to wave action
Journal of Experimental Biology 216:1717-1725 (DOI: 10.1242/jeb.078964).
(download from journal or request pdf from me)
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Palmer, A.R. 2012.
Developmental plasticity and the origin of novel forms:
Unveiling cryptic genetic variation via "use and disuse"
Journal of Experimental Zoology (Molecular & Developmental Evolution) 318:466Ð479.
(download from journal or request pdf from me)
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Palmer, A.R. 2012.
Developmental origins of normal and anomalous random right-left asymmetry:
Lateral inhibition versus developmental error in a threshold trait
Contributions to Zoology 81(2):111-124.
(download from journal or request pdf from me)
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Neufeld, C.* & A.R. Palmer*. 2011.
Learning, developmental plasticity, and the rate of morphological evolution. p. 337-356.
In: Epigenetics: Linking Genotype and Phenotype in Development and Evolution (B. Hallgrimmson & B.K. Hall, eds.) Berkeley, California. University of California Press.
(order book from publisher or request pdf from me.)
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Palmer, A.R. (translator) 2011.
History of a mussel bed. Observations on a phase of faunal disequilibrium (English translation of: Fischer-Piette, E. 1935. Histoire d'une Mouliere. Observations sur une phase de desequilibre faunique. Bulletin Biologique de la France et de la Belgique 5:153-177).
In: Essential Naturalist: Timeless Readings in Natural History (Graham, M.H., J. Parker & P.K. Dayton, eds.) University of Chicago Press, Chicago.
(download preprint: 177K)
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(66 cites)
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Neinhuis, S.*, A.R. Palmer* & C. Harley. 2010.
Elevated CO2 affects shell dissolution rate but not calcification rate in a marine snail.
Proceedings of the Royal Society of London B 277: 2553-2558.
(download from journal or request pdf from me)
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Hinchliffe, R.* & A.R. Palmer*. 2010.
Curious coiled cases of caddisfly larvae: Handed behavior, ontogeny and evolution.
Integrative Biology 50(4): 606-618.
(download from journal or request pdf from me)
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MEDIA
COVERAGE:
Journal
Cover
Neue Zurcher
Zeitung(11/2010)
Die Aquarienzt-
schrift (01/2011)
Illustreret Viden-
skab (05/2011)
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Krause-Nehring, J., M.J. Starck & A.R. Palmer. 2010.
Juvenile colour polymorphism in the red rock crab, Cancer productus: Patterns, causes, and possible adaptive significance.
Zoology 113: 131-139.
(download from journal or request pdf from me)
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Palmer, A.R. 2010.
Scale-eating cichlids: From hand(ed) to mouth.
Journal of Biology 9(2):11.
(download from journal- Open Access)
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Rosen, E., B. Schwarz & A.R. Palmer. 2009.
Smelling the difference: Hermit crab responses to predatory and non-predatory crabs.
Animal Behaviour 78: 691-695.
(download from journal or request pdf from me)
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Edgell, T.C., B.R. Lynch, G.C. Trussell & A.R. Palmer. 2009.
Experimental evidence for the rapid evolution of behavioral canalization in natural populations.
American Naturalist 174 (2): 434-440.
(download from journal or request pdf from me)
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MEDIA
COVERAGE
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Neufeld, C. & A.R. Palmer. 2008.
Precisely proportioned: Intertidal barnacles alter penis form to suit coastal wave action.
Proceedings of the Royal Society of London. B. 275:1081-1087.
(download from journal or request pdf from me)
(movie of barnacles copulating)
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Bergstrom, C.A. & A.R. Palmer. 2007.
Which way to turn? Effect of direction of body asymmetry on turning and prey strike orientation in starry flounder (Platichthys stellatus, Pleuronectidae).
Journal of Fish Biology 71: 737-748.
(download: 393K)
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Rypien, K.L. & A.R. Palmer. 2007.
The effect of sex, size and habitat on the incidence of puncture wounds in the claws of the porcelain crab Petrolisthes cinctipes (Anomura: Porcellanidae).
Journal of Crustacean Biology 27: 59-64.
(download: 140K)
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(95 cites)
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Levin, M. & A.R. Palmer. 2007.
Left-right patterning from the inside out: Widespread evidence for intracellular control.
BioEssays 29: 271-287.
(download: 1.1MB)
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Palmer, A.R. 2006.
Caught right-handed. (News & Views)
Nature 444: 689-691.
(download: 604K)
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(90 cites)
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Anker, A., S.T. Ahyong, P.Y. Noel & A.R. Palmer. 2006.
Morphological phylogeny of alpheid shrimps: Parallel preadaptation and the origin of a key morphological innovation - the snapping claw.
Evolution 60: 2507-2528.
(download; includes supplementary information: 4.9MB)
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Marin, I.N., A. Anker, T.A. Britayev & A.R. Palmer. 2005.
Symbiosis between the alpheid shrimp Athanas ornithorhynchus Banner and Banner, 1973 (Crustacea: Decapoda), and the brittle star Macrophiothrix longipeda (Lamarck, 1816) (Echinodermata: Ophiuroidea)
Zoological Studies 44:234-241.
(download: 586K)
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Anker, A., G.-V. Murina, C. Lira, J.A. Vera Caripe, A.R. Palmer & M.-S. Jeng. 2005.
Macrofauna associated with echiuran burrows: A review with new observations on the innkeeper worm Ochetostoma erythrogrammon Leuckart and Ruppel, in Venezuela.
Zoological Studies 44:157-192.
(download: 1.5MB)
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Palmer, A.R. 2005.
Is the ID debate proof of an intelligent deceiver? (letter to editor)
Nature 434:422.
(download: 86K)
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Palmer, A.R. (posted 2005)
Irregularities in data reported by A.P. Moller
(unpublished MS)
(download: 128K)
(view entire exchange regarding this document on the EvolDir listserv)
Additional questions have also been raised about questionable data from, or conduct by, Anders Moller.
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Palmer, A.R. 2005.
Antisymmetry. pp. 359-397
In: Variation (B. Hallgrimmson & B. K. Hall, eds) Elsevier (order the book or request pdf from me)
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Palmer, A.R. 2004.
Selection for asymmetry. (letter to editor)
Science 306:812.
(download from journal or request pdf from me)
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MEDIA
COVERAGE:
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Eaves, A.A. & Palmer, A.R. 2003.
Widespread cloning in echinoderm larvae.
Nature 425:146.
(download: 83K)
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Palmer, A.R. 2003.
Reply to Hopkins and Cantalupo: Chimpanzee right-handedness reconsidered - Sampling issues and data presentation.
American Journal of Physical Anthropology 121:382-384.
(download: 55K)
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Journal
Cover
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Marchinko, K.B. & Palmer, A.R. 2003.
Feeding in flow extremes: Dependence of cirrus form on wave-exposure in four barnacle species.
Zoology 106:127-141.
(download: 1,156K)
(download electronic Appendix: 42K)
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(337 cites)
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Palmer, A.R. & C. Strobeck. 2003.
Fluctuating asymmetry analyses revisited.
In Developmental Instability (DI): Causes and Consequences,
M. Polak, ed. Oxford University Press, Oxford. pp. 279-319 (the book may be ordered here).
(download preprint, 236K; download electronic Appendix V, 103K; download data files)
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Palmer, A.R. 2002.
Chimpanzee right-handedness reconsidered: Evaluating the evidence with funnel plots.
American Journal of Physical Anthropology 118:191-199.
(download: 177K)
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Arsenault, D.J., K.B. Marchinko, & A.R. Palmer. 2001.
Precise tuning of barnacle leg length to coastal wave action.
Proceedings of the Royal Society of London. B 268:2149-2154.
(download: 270K)
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Palmer, A.R. & L.M. Hammond. 2000.
The emperor's codpiece: A postmodern perspective on biological asymmetries.
International Society for Behavioral Ecology, Newsletter 12:13-20.
(download: 104K)
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Taylor, G.M., A.R. Palmer & A.C. Barton. 2000.
Variation in safety factors of claws within and among six species of Cancer crabs (Decapoda: Brachyura).
Biological Journal of the Linnean Society 70: 37-62.
(download: 251K)
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Palmer, A.R., G.M. Taylor & A.C. Barton. 1999.
Cuticle strength and the size-dependence of safety factors in Cancer crab claws.
Biological Bulletin 196:281-294.
(download: 1549K)
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(206 cites)
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Palmer, A.R. 1999.
Detecting publication bias in meta-analyses: A case study of fluctuating asymmetry and sexual selection.
American Naturalist 154: 220-233.
(download: 269K)
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Palmer, A.R. 1994.
Temperature sensitivity, rate of development and generation time: Geographic variation in laboratory-reared Nucella and a cross-phyletic overview.
In: Reproduction and Development of Marine Invertebrates. (W.H. Wilson, S.A. Stricker, G.L. Shinn, Eds.). Johns Hopkins University Press, Baltimore pp. 177-194.
(download: 616K)
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Chippindale, A.K.. & A.R. Palmer. 1993. Persistence of statistical departures from symmetry over multiple molts in individual brachyuran crabs: Relevance to developmental stability.
Genetica 89:185-199.
(download: 1.2MB)
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Palmer, A.R. 1992.
Intraspecific variation in three species of rocky shore gastropods from Hong Kong: Correlations among habitats and a comparison with temperate species.
In: Proceedings of the Fourth International Marine Biological Workshop: The Marine Flora & Fauna of Hong Kong and Southern China. Vol. 1. General Proceedings. (B. Morton, Ed.). Hong Kong University Press pp. 649-691.
(download: 1.4MB)
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Palmer, A.R. S.D. Gayron & D.S. Woodruff. 1990.
Reproductive, morphological, and genetic evidence for two cryptic species of Northeastern Pacific Nucella.
Veliger 33:325-338.
(download: 605K)
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Vermeij, G.J., A.R. Palmer & D.R. Lindberg. 1990.
Range limits and dispersal of molluscs in the Aleutian Islands, Alaska.
Veliger 33:346-354.
(download: 437K)
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(298 cites)
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Appleton, R.D.* & A.R. Palmer*. 1988.
Water-borne stimuli released by predatory crabs and damaged prey induce more predator-resistant shells in a marine gastropod.
Proceedings of the National Academy of Sciences (USA) 85:4387-4391.
(download: low resolution, 305K;
high resolution, 1.4MB)
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Palmer, A.R. 1988.
Feeding biology of Ocenebra lurida (Prosobranchia:Muricacea): Diet, predator-prey size relations, and attack behavior.
Veliger 31:192-203.
(download: 625K)
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Hart, M.W. & A.R. Palmer 1987.
Stereotypy, ontogeny and heritability of drill site selection in thaidid gastropods.
Journal of Experimental Marine Biology and Ecology 107:101-120.
(download: 1.7MB)
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(1515 cites)
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Palmer, A.R. & C. Strobeck. 1986.
Fluctuating asymmetry: Measurement, analysis, patterns.
Annual Review of Ecology and Systematics 17:391-421.
(download: low resolution, 737K;
high resolution, 2.6MB)
[by 2009, this paper was the 9th most highly cited paper of 767 ever published in ARES]
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Palmer, A.R. 1985.
Quantum changes in gastropod shell morphology need not reflect speciation.
Evolution 39(3):699-705.
(download: low resolution, 259K;
high resolution, 933K)
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Mercurio, K.S., R.B. Lowell & A.R. Palmer 1985.
Predator-mediated microhabitat partitioning by two species of visually cryptic, intertidal limpets.
Ecology 66:1417-1425.
(download: 386K)
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Palmer, A.R. 1985.
Genetic basis of shell variation in Thais emarginata (Prosobranchia, Muricacea) I. Banding in populations from Vancouver Island.
[now known as Nucella ostrina]
Biological Bulletin 169:638-651.
(download: 386K)
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Palmer, A.R. 1985.
Adaptive value of shell variation in Thais (or Nucella) lamellosa: Effect of thick shells on vulnerability to and preference by crabs.
Veliger 27:349-356.
(download: 339K)
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Palmer, A.R. 1984.
Prey selection by thaidid gastropods: Some field tests of foraging models.
Oecologia 62:162-172.
(download: 2.0MB)
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Palmer, A.R. 1984.
Species cohesiveness and genetic control of shell color and form in Thais emarginata (Prosobranchia, Muricacea): Preliminary results.
[now known as Nucella ostrina]
Malacologia 25:477-491.
(download: 490K)
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Palmer, A.R. 1983.
Relative costs of producing skeletal organic matrix versus calcification: Evidence from marine gastropods.
Marine Biology 75:287-292.
(download: 2.3MB)
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Palmer, A.R. 1983.
Growth rate as a measure of food value in thaidid gastropods: Assumptions and implications for prey morphology and distribution.
Journal of Experimental Marine Biology and Ecology 73:95-124.
(download: 2.1MB)
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Palmer, A.R. 1982.
Predation and parallel evolution: Recurrent parietal plate reduction in balanomorph barnacles.
Paleobiology 8:31-44.
(download: low resolution, 522K;
high resolution, 1.9MB)
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Palmer, A.R. J. Szymanska & L. Thomas 1982.
Prolonged withdrawal: A possible predator evasion behavior in Balanus glandula (Crustacea, Cirripedia).
Marine Biology 67:51-55.
(download: 509K)
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Palmer, A.R. 1982.
Growth in marine gastropods: A non-destructive technique for independently measuring shell and body weight.
Malacologia 23:63-73.
(download: 387K)
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(125 cites)
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Palmer, A.R. 1981.
Do carbonate skeletons limit the rate of body growth?
Nature 292:150-152.
(download: 325K)
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Palmer, A.R. 1981.
Predator errors, foraging in unpredictable environments and risk: The consequences of prey variation in handling time versus net energy
American Naturalist 118:908-915.
(download: 765K)
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(129 cites)
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Palmer, A.R. & R.R. Strathmann 1981.
Scale of dispersal in varying environments and its implications for life histories of marine invertebrates.
Oecologia 48:308-318.
(download: 1.3MB)
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Palmer, A.R. 1980.
Locomotion rates and shell form in the Gastropoda: A re-evaluation.
Malacologia 19:289-296.
(download: 215K)
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(238 cites)
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Palmer, A.R. 1979.
Fish predation and the evolution of gastropod shell sculpture: Experimental and geographic evidence.
Evolution 33:697-713.
(download: low resolution, 642K;
high resolution, 2.8MB)
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Paine, R.T. & A.R. Palmer 1978.
Sicyases sanguineus: A unique trophic generalist from the Chilean intertidal zone.
Copeia 1978:75-81.
(download: 520K)
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Palmer, A.R. 1977.
Function of shell sculpture in marine gastropods: Hydrodynamic destabilization in Ceratostoma foliatum.
Science 197:1293-1295.
(download: low resolution, 228K;
high resolution, 1.2MB)
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Palmer, A.R. 2000.
Invertebrate Zoology teaching animations: Animations of form and function in many invertebrates, including choanoflagellate protists, sponges, hydroids, jellyfish, bilaterian excretory systems, filibranch and eulamellibranch bivalves, polychaete and oligochaete annelid worms, brachyuran crab claws, sea urchins, and sea squirts. A cool Invertebrate diversity quiz is a bonus.
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Palmer, A.R. 2021.
Guess the Invertebrate: A special on-line game created for the 25th anniversary edition of the journal Invertebrate Biology.
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Palmer, A.R. 2026.
MacCoil (Javascript version): Displays wire-frame images of coiled mollusc shells drawn using user-specified ('point-and-click') coiling parameters in any browser.
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NOTE: these programs only run on legacy Macintosh computers (those that can run in "Classic" mode or earlier).
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Palmer, A.R. 1992, 1994.
ExCaliper: Captures data from digital calipers onto a Macintosh computer.
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Palmer, A.R. 1996.
MacCoil (Pascal version): Displays wire-frame images of coiled mollusc shells drawn using user-specified ('point-and-click') coiling parameters on a Macintosh computer.
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- T.H. Huxley's 'The Crayfish' on-line (a WONDERFUL, turn-of-the-century essay on Zoology for the layman; complete with hot-links to a glossary and copies of the original woodcut figures)
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